I am interested in learning how the Arduino IDE handles parameters in order to correctly upload hex files with avrdude.

I found this in boards.txt:

uno.name=Arduino/Genuino Uno

uno.vid.0=0x2341
uno.pid.0=0x0043
uno.vid.1=0x2341
uno.pid.1=0x0001
uno.vid.2=0x2A03
uno.pid.2=0x0043
uno.vid.3=0x2341
uno.pid.3=0x0243

uno.upload.tool=avrdude
uno.upload.protocol=arduino
uno.upload.maximum_size=32256
uno.upload.maximum_data_size=2048
uno.upload.speed=115200

uno.bootloader.tool=avrdude
uno.bootloader.low_fuses=0xFF
uno.bootloader.high_fuses=0xDE
uno.bootloader.extended_fuses=0x05
uno.bootloader.unlock_bits=0x3F
uno.bootloader.lock_bits=0x0F
uno.bootloader.file=optiboot/optiboot_atmega328.hex

uno.build.mcu=atmega328p
uno.build.f_cpu=16000000L
uno.build.board=AVR_UNO
uno.build.core=arduino
uno.build.variant=standard

There is this in platform.txt:

# AVR Uploader/Programmers tools
# ------------------------------

tools.avrdude.path={runtime.tools.avrdude.path}
tools.avrdude.cmd.path={path}/bin/avrdude
tools.avrdude.config.path={path}/etc/avrdude.conf

tools.avrdude.upload.params.verbose=-v
tools.avrdude.upload.params.quiet=-q -q
tools.avrdude.upload.pattern="{cmd.path}" "-C{config.path}" {upload.verbose} -p{build.mcu} -c{upload.protocol} -P{serial.port} -b{upload.speed} -D "-Uflash:w:{build.path}/{build.project_name}.hex:i"

tools.avrdude.program.params.verbose=-v
tools.avrdude.program.params.quiet=-q -q
tools.avrdude.program.pattern="{cmd.path}" "-C{config.path}" {program.verbose} -p{build.mcu} -c{protocol} {program.extra_params} "-Uflash:w:{build.path}/{build.project_name}.hex:i"

tools.avrdude.erase.params.verbose=-v
tools.avrdude.erase.params.quiet=-q -q
tools.avrdude.erase.pattern="{cmd.path}" "-C{config.path}" {erase.verbose} -p{build.mcu} -c{protocol} {program.extra_params} -e -Ulock:w:{bootloader.unlock_bits}:m -Uefuse:w:{bootloader.extended_fuses}:m -Uhfuse:w:{bootloader.high_fuses}:m -Ulfuse:w:{bootloader.low_fuses}:m

tools.avrdude.bootloader.params.verbose=-v
tools.avrdude.bootloader.params.quiet=-q -q
tools.avrdude.bootloader.pattern="{cmd.path}" "-C{config.path}" {bootloader.verbose} -p{build.mcu} -c{protocol} {program.extra_params} "-Uflash:w:{runtime.platform.path}/bootloaders/{bootloader.file}:i" -Ulock:w:{bootloader.lock_bits}:m

This line is particularly interesting:

tools.avrdude.upload.pattern="{cmd.path}" "-C{config.path}" {upload.verbose} -p{build.mcu} -c{upload.protocol} -P{serial.port} -b{upload.speed} -D "-Uflash:w:{build.path}/{build.project_name}.hex:i"

It seems like these variables are corresponding:

{cmd.path} --> tools.avrdude.cmd.path
{config.path} --> tools.avrdude.config.path
{upload.verbose} --> tools.avrdude.upload.params.verbose
{build.mcu} --> uno.build.mcu
{upload.protocol} --> uno.upload.protocol
{serial.port} --> ?
{upload.speed} --> uno.upload.speed
{build.path} --> ?
{build.project_name} --> ?

It seems these are likely used in the Java code for the IDE. There are obviously a few I am unsure of. I would guess these are considered user inputs in the IDE. Though I am by no means a Java or Javascript programmer.

Where are these variables explicitly defined in the Arduino library? (e.g. {runtime.tools.avrdude.path}, {path}, {cmd.path} ....) Or how do they get defined by IDE?